The Belgian took an emotional victory, which he dedicated to Fabio Jakobsen, after attacking with 51 kilometers to go.
At the end of 305 kilometers, the longest distance in the history of the race, the defending champion lost for a matter of centimeters.
A consistent ride on the tough stage 2 helped the 22-year-old Portuguese cement his place in the top 10 overall.
The 20-year-old US rider will compete as a stagiaire for the Wolfpack.
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